Fiddler Crabs Home

Thurman, C.L., II, J.C. McNamara, H.-T. Shih, and M.V. Capparelli (2025) Fiddler crabs (Crustacea: Decapoda: Ocypodidae) from coastal Ecuador and the Galápagos Islands: Species descriptions and DNA barcodes. Ecology and Evolution 15(1):e70646.

Rio Coyol Rio Coyol

Language: English

Online: DOI https://doi.org/10.1002/ece3.70646

Names Appearing in this Publication

Name Used Common Name(s) Where Applied to... Accepted Name Source of Accepted Note(s)
  fiddler crabs text p. 1-37 citation: Aoki & Wada (2013) TBD Computed  
      citation: Barnwell & Thurman (1984) TBD Computed  
      citation: Barnwell & Szelistowski (1985) “fiddler crabs” Original  
      citation: Crane (1941) “fiddler crabs” Original  
      citation: Crane (1975)Uca “fiddler crabs” Computed in part
      citation: Hardianto et al. (2022) TBD Computed  
      citation: Hendrickx (1979) “fiddler crabs” Original  
      citation: Hendrickx (1995) “fiddler crabs” Original  
      citation: Latreille (1817)Gelasima Genus Gelasimus Computed  
      citation: Laurenzano et al. (2012) TBD Computed  
      citation: Laurenzano et al. (2016) TBD Computed  
      citation: Lemaitre & Alvarez León (1992)Uca “fiddler crabs” Computed  
      citation: Marochi et al. (2022) TBD Computed  
      citation: Nehemia & Kochzius (2017) TBD Computed  
      citation: Rathbun (1897)Uca Genus Uca Computed in part
      citation: Rathbun (1935) “fiddler crabs” Original  
      citation: Rosenberg (2020). “fiddler crabs” Computed in part
      citation: Shih et al. (2015) TBD Computed  
      citation: Shih et al. (2016) “fiddler crabs” Original  
      citation: Shih et al. (2022). “fiddler crabs” Computed in part
      citation: Shih et al. (2022) TBD Computed  
      citation: Shih & Chan (2022) TBD Computed  
      citation: Silva et al. (2010) TBD Computed  
      citation: Staton et al. (2014) TBD Computed  
      citation: Thurman (2002) TBD Computed  
      citation: Thurman (2003) TBD Computed  
      citation: Thurman (2003) TBD Computed  
      citation: Thurman et al. (2010) TBD Computed  
      citation: Thurman et al. (2013) TBD Computed  
      citation: Thurman et al. (2018) TBD Computed  
      citation: Thurman et al. (2021) TBD Computed  
      citation: Thurman et al. (2023) TBD Computed  
      citation: Tokuyama et al. (2020) TBD Computed  
      citation: Wieman et al. (2014) TBD Computed  
      citation: von Hagen (1968) “fiddler crabs” Original  
      citation: von Prahl (1982)Uca “fiddler crabs” Computed  
      citation: von Prahl & Toro (1985) “fiddler crabs” Original  
      location: Ecuador “fiddler crabs” Original  
      location: El Salvador “fiddler crabs” Original  
      location: Galápagos Islands “fiddler crabs” Original  
      location: Pacific coast of Colombia “fiddler crabs” Original  
      location: Pacific coast of Panama “fiddler crabs” Original  
 
Austruca citrus   text p. 35 citation: Shih & Poupin (2020)Austruca citrus Austruca citrus Computed in part
 
Austruca occidentalis   text p. 33 citation: Naderloo et al. (2016)Uca occidentalis Austruca annulipes Computed in part
 
Austruca perplexa   text p. 35 citation: Shih & Poupin (2020)Austruca perplexa Austruca perplexa Computed in part
 
Austruca variegata   text p. 35 citation: Shih et al. (2019)Austruca variegata Austruca variegata Computed in part
 
Gelasimus hesperiae   text p. 35 citation: Shih et al. (2010)Uca hesperiae Gelasimus hesperiae Computed  
 
Leptuca   text p. 1, 4-5, 8-9, 13, 21, 32, 36 citation: Shih et al. (2016)Leptuca Genus Leptuca Computed in part
      location: Ecuador Genus Leptuca Original  
 
Leptuca aff. dorotheae   text p. 8, 34 specimen: Rio Tempisque, Guanacaste, Costa Rica [ZRC] Minuca zacae Original  
 
Leptuca batuenta   text p. 2-3, 5, 8, 13, 21-22, 34, 37; Figure 14; Supplement p. 6 citation: von Hagen (1968)Uca batuenta Leptuca batuenta Computed  
      specimen: Balboa, Panama City, Panama [USNM 137405] Leptuca batuenta Original  
      specimen: Balboa, Panama City, Panama [USNM 137406] Leptuca batuenta Original  
      specimen: Ballena Bay, Puntarenas, Costa Rica [USNM 137404] Leptuca batuenta Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 861] Leptuca batuenta Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138473] Leptuca batuenta Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23045] Leptuca batuenta Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376053] Leptuca batuenta Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376056] Leptuca batuenta Original  
 
Leptuca beebei   text p. 2-3, 5, 8, 13, 21-22, 32-35, 37; Figure 15; Supplement p. 6 citation: Crane (1975)Uca (Celuca) beebei Leptuca beebei Computed  
      citation: von Hagen (1968)Uca beebei Leptuca beebei Computed  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI 764] Leptuca beebei Original  
      specimen: Balboa, Panama City, Panama [USNM 137413] Leptuca beebei Original  
      specimen: Balboa, Panama City, Panama [USNM 137414] Leptuca beebei Original  
      specimen: Balboa, Panama City, Panama [USNM 79400] Leptuca beebei Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376086] Leptuca beebei Original  
      specimen: Guerrero, Mexico [AMNH_IZC 376060] Leptuca beebei Original  
      specimen: Gulf of Fonseca, La Unión, El Salvador [RMNH.CRUS.D9499] Leptuca beebei Original  
      specimen: Manzanillo, Colima, Mexico [AMNH_IZC 376059] Leptuca beebei Original  
      specimen: Posorja, Guayas, Ecuador [UNI 763] Leptuca beebei Original  
      specimen: Posorja, Guayas, Ecuador [UNI 793] Leptuca beebei Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [UNI 794] Leptuca beebei Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138480] Leptuca beebei Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23056] Leptuca beebei Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376061] Leptuca beebei Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 865] Leptuca beebei Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI 762] Leptuca beebei Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376070] Leptuca beebei Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 791] Leptuca beebei Original  
      specimen: Usulután, El Salvador [SMF 48165] Leptuca beebei Original  
 
Leptuca deichmanni   text p. 2-3, 5, 8, 13, 22-23, 33-34, 37; Figure 16; Supplement p. 6-7 citation: von Prahl (1982)Uca (Celuca) deichmanni Leptuca deichmanni Computed  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI 890] Leptuca deichmanni Original  
      specimen: Bahía de Santa Elena, Guanacaste, Costa Rica [USNM 138521] Leptuca deichmanni Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 138524] Leptuca deichmanni Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376108] Leptuca deichmanni Original  
      specimen: Honda Bay, Panama [USNM 138525] Leptuca deichmanni Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13583] Leptuca deichmanni Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 792] Leptuca deichmanni Original  
      specimen: Panama Canal, Panama [USNM 70832] Leptuca deichmanni Original  
      specimen: Panama City, Panama [USNM 138526] Leptuca deichmanni Original  
      specimen: Playa Achilube, Esmeraldas, Ecuador [UNI 765] Leptuca deichmanni Original  
      specimen: Playa Achilube, Esmeraldas, Ecuador [UNI P0050] Leptuca deichmanni Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376104] Leptuca deichmanni Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376105] Leptuca deichmanni Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 766] Leptuca deichmanni Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 868] Leptuca deichmanni Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI 767] Leptuca deichmanni Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376108] Leptuca deichmanni Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 772] Leptuca deichmanni Original  
 
Leptuca dorotheae   text p. 2-3, 5-6, 8, 13, 23-24, 33-34, 37; Figure 17; Supplement p. 7 citation: Rosenberg (2020)Leptuca dorotheae Leptuca dorotheae Computed  
      citation: von Hagen (1968)Uca dorotheae Leptuca dorotheae Computed  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI P0306] Leptuca dorotheae Original  
      specimen: Balboa, Panama City, Panama [USNM 138534] Leptuca dorotheae Original  
      specimen: Puento Carnero, Santa Elena, Ecuador [UNI 773] Leptuca dorotheae Original  
      specimen: Puento Carnero, Santa Elena, Ecuador [UNI P0271] Leptuca dorotheae Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138534] Leptuca dorotheae Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23054] Leptuca dorotheae Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23055] Leptuca dorotheae Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376113] Leptuca dorotheae Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376114] Leptuca dorotheae Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 771] Leptuca dorotheae Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI P0269] Leptuca dorotheae Original  
 
Leptuca festae   text p. 2-3, 5, 8, 13, 24, 34, 37; Figure 18; Supplement p. 7 citation: Crane (1975)Uca (Celuca) festae Leptuca festae Computed  
      citation: Rathbun (1918)Uca festae Leptuca festae Computed  
      citation: Rathbun (1935)Uca guayaquilensis Leptuca festae Computed  
      citation: Shih et al. (2016)Leptuca festae Leptuca festae Computed in part
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 770] Leptuca festae Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376118] Leptuca festae Original  
      specimen: Guayaquil, Guayas, Ecuador [USNM 138533] Leptuca festae Original  
      specimen: Guayaquil, Guayas, Ecuador [USNM 70870] Leptuca festae Original  
      specimen: Los Lojas, Guayaquil, Guayas, Ecuador [UNI P0212] Leptuca festae Original  
      specimen: Puento Montalvo, Esmeraldas, Ecuador [UNI 775] Leptuca festae Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376115] Leptuca festae Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 776] Leptuca festae Original  
      specimen: Rio Esmeraldas, Esmeraldas, Ecuador [UNI 774] Leptuca festae Original  
      specimen: Rio Guayas, Guayas, Ecuador [UNI 751] Leptuca festae Original  
 
Leptuca helleri   text p. 2-3, 5, 8, 13, 24-25, 33-34, 37; Figure 19; Supplement p. 7 citation: Capparelli et al. (2021) TBD Computed  
      citation: Rathbun (1918)Uca helleri Leptuca helleri Computed  
      specimen: Fernandina Island, Galápagos Islands [USNM 24829] Leptuca helleri Original  
      specimen: Fernandina Island, Galápagos Islands [USNM 25666] Leptuca helleri Original  
      specimen: Isabela Island, Galápagos Islands [USNM 63154] Leptuca helleri Original  
      specimen: Santa Cruz Island, Galápagos Islands [UNI 777] Leptuca helleri Original  
      specimen: Santa Cruz Island, Galápagos Islands [UNI P0102] Leptuca helleri Original  
      specimen: Santa Cruz Island, Galápagos Islands [UNI P0103] Leptuca helleri Original  
      specimen: Tortuga Bay, Santa Cruz Island, Galápagos Islands [UNI 778] Leptuca helleri Original  
 
Leptuca inaequalis   text p. 3, 5, 8, 13, 25-26, 34, 37; Figure 20; Supplement p. 7 specimen: El Rompido, Esmeraldas, Ecuador [UNI 780] Leptuca inaequalis Original  
      specimen: Estero Salado, Guayaquil, Guayas, Ecuador [USNM 138552] Leptuca inaequalis Original  
      specimen: Estero Salado, Guayaquil, Guayas, Ecuador [USNM 70833] Leptuca inaequalis Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376140] Leptuca inaequalis Original  
      specimen: Puenta Sitio Nuevo, Santa Elena, Ecuador [UNI 782] Leptuca inaequalis Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138551] Leptuca inaequalis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23043] Leptuca inaequalis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376132] Leptuca inaequalis Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 781] Leptuca inaequalis Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 871] Leptuca inaequalis Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI 779] Leptuca inaequalis Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI P0273] Leptuca inaequalis Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI P0274] Leptuca inaequalis Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376138] Leptuca inaequalis Original  
 
Leptuca latimanus   text p. 2-3, 5, 9, 13, 25-26,32-34, 37; Figure 21; Supplement p. 7-8 citation: Crane (1975)Uca (Celuca) latimanus Leptuca latimanus Computed  
      citation: Rathbun (1918)Uca latimanus Leptuca latimanus Computed in part
      specimen: Balboa, Panama City, Panama [USNM 138565] Leptuca latimanus Original  
      specimen: El Real, Santa Elena, Ecuador [UNI P0152] Leptuca latimanus Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376146] Leptuca latimanus Original  
      specimen: La Paz, Baja California Sur, Mexico [USNM 17500] Leptuca latimanus Original  
      specimen: Manglaraltos, Santa Elena, Ecuador [UNI 788] Leptuca latimanus Original  
      specimen: Nayarit, Mexico [AMNH_IZC 376143] Leptuca latimanus Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376148] Leptuca latimanus Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376154] Leptuca latimanus Original  
      specimen: Playa Camarones, Esmeraldas, Ecuador [UNI 787] Leptuca latimanus Original  
      specimen: Posorja, Guayas, Ecuador [UNI 789] Leptuca latimanus Original  
      specimen: Puenta Las Javitas, Santa Elena, Ecuador [UNI 784] Leptuca latimanus Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138566] Leptuca latimanus Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23052] Leptuca latimanus Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376149] Leptuca latimanus Original  
      specimen: Rio Cadente, Santa Elena, Ecuador [UNI 785] Leptuca latimanus Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 783] Leptuca latimanus Original  
 
Leptuca limicola   text p. 5, 33 citation: Rosenberg (2020)Leptuca limicola Leptuca limicola Computed  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137415] Leptuca limicola Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137416] Leptuca limicola Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 79401] Leptuca limicola Original  
 
Leptuca oerstedi   text p. 33 citation: Rosenberg (2020)Leptuca oerstedi Leptuca oerstedi Computed  
 
Leptuca pygmaea   Supplement p. 8 specimen: Bahia de Buenaventura, Valle del Cauca, Colombia [USNM 138646] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137419] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137420] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Panama Canal, Panama [USNM 1521269] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376256] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: San Lorenzo, Honduras [AMNH_IZC 376258] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
 
Leptuca saltitanta   text p. 3, 5, 9, 13, 26-27, 33-34, 37; Figure 22; Supplement p. 8 citation: Crane (1975)Uca (Celuca) [crenulata] saltitanta Leptuca saltitanta Computed  
      specimen: Balboa, Panama City, Panama [USNM 137407] Leptuca saltitanta Original  
      specimen: Balboa, Panama City, Panama [USNM 137408] Leptuca saltitanta Original  
      specimen: Balboa, Panama City, Panama [USNM 138822] Leptuca saltitanta Original  
      specimen: Balboa, Panama City, Panama [USNM 79403] Leptuca saltitanta Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 795] Leptuca saltitanta Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI P0282] Leptuca saltitanta Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 866] Leptuca saltitanta Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376260] Leptuca saltitanta Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI 869] Leptuca saltitanta Original  
 
Leptuca sp.   text p. 9, 24, 33-34 citation: Crane (1975) ? Original Reference to an unidentified specimen
      specimen: Esmeraldas, Ecuador [UNI P0289] ? Original  
      specimen: Esmeraldas, Ecuador [UNI P0290] ? Original  
 
Leptuca stenodactylus   text p. 2-3, 6, 9, 13, 27-28, 32-35, 37; Figure 23; Supplement p. 8 citation: Crane (1975)Uca (Celuca) stenodactylus Leptuca stenodactylus Computed  
      citation: Rathbun (1918)Uca stenodactylus Leptuca stenodactylus Computed in part
      specimen: Corinto, Nicaragua [USNM 138826] Leptuca stenodactylus Original  
      specimen: Costa Rica [USNM 138830] Leptuca stenodactylus Original  
      specimen: El Salvador [SMF 2357] Leptuca stenodactylus Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376319] Leptuca stenodactylus Original  
      specimen: Panama City, Panama [USNM 138830] Leptuca stenodactylus Original  
      specimen: Posorja, Guayas, Ecuador [UNI 749] Leptuca stenodactylus Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138833] Leptuca stenodactylus Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23057] Leptuca stenodactylus Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376317] Leptuca stenodactylus Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 769] Leptuca stenodactylus Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 768] Leptuca stenodactylus Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376311] Leptuca stenodactylus Original  
 
Leptuca tallanica   text p. 2-3, 5-7, 9, 13, 28-29, 32-34, 37; Figure 24; Supplement p. 8 citation: Rosenberg (2020)Leptuca tallanica Leptuca tallanica Computed  
      citation: von Hagen (1968)Uca tallanica Leptuca tallanica Computed  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 862] Leptuca tallanica Original  
      specimen: Machala, El Oro, Ecuador [UNI P0280] Leptuca tallanica Original  
      specimen: Machala, El Oro, Ecuador [UNI P279] Leptuca tallanica Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 863] Leptuca tallanica Original  
      specimen: Posorja, Guayas, Ecuador [UNI P277] Leptuca tallanica Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138838] Leptuca tallanica Original  
      specimen: Puerto El Morro, Guayas, Ecuador [UNI P0117] Leptuca tallanica Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23044] Leptuca tallanica Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23046] Leptuca tallanica Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23047] Leptuca tallanica Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 867] Leptuca tallanica Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 864] Leptuca tallanica Original  
 
Leptuca tenuipedis   text p. 2-3, 5-7, 13, 28-30, 37; Figure 25; Supplement p. 8-9 citation: von Hagen (1968)Uca tenuipedis Leptuca tenuipedis Computed  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI 891] Leptuca tenuipedis Original  
      specimen: Ballena Bay, Puntarenas, Costa Rica [USNM 137409] Leptuca tenuipedis Original  
      specimen: Ballena Bay, Puntarenas, Costa Rica [USNM 137410] Leptuca tenuipedis Original  
      specimen: Ballena Bay, Puntarenas, Costa Rica [USNM 79404] Leptuca tenuipedis Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376330] Leptuca tenuipedis Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 866] Leptuca tenuipedis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23044] Leptuca tenuipedis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376331] Leptuca tenuipedis Original  
 
Leptuca terpsichores   text p. 2-3, 5, 9, 13, 30, 32-34, 37; Figure 26; Supplement p. 9 citation: Crane (1975)Uca (Celuca) musica terpsichores Leptuca terpsichores Computed  
      citation: von Hagen (1968)Uca terpsichores Leptuca terpsichores Computed  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 796] Leptuca terpsichores Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376372] Leptuca terpsichores Original  
      specimen: Isla Jambeli, El Oro, Ecuador [USNM 138608] Leptuca terpsichores Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13582] Leptuca terpsichores Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 797] Leptuca terpsichores Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 800] Leptuca terpsichores Original  
      specimen: Panama Canal, Panama [USNM 137417] Leptuca terpsichores Original  
      specimen: Panama Canal, Panama [USNM 137418] Leptuca terpsichores Original  
      specimen: Puento Carnero, Santa Elena, Ecuador [UNI 799] Leptuca terpsichores Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23053] Leptuca terpsichores Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376335] Leptuca terpsichores Original  
      specimen: San Lorenzo Malecón, Esmeraldas, Ecuador [UNI 798] Leptuca terpsichores Original  
 
Leptuca tomentosa   text p. 2-3, 6, 9, 13, 30-31, 34-35, 37; Figure 27; Supplement p. 9 citation: Crane (1975)Uca (Celuca) tomentosa Leptuca tomentosa Computed  
      citation: von Hagen (1968)Uca mertensi Leptuca tomentosa Computed  
      specimen: El Rompido, Esmeraldas, Ecuador [USM 803] Leptuca tomentosa Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 138839] Leptuca tomentosa Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376398] Leptuca tomentosa Original  
      specimen: Muisne, Esmeraldas, Ecuador [USM 801] Leptuca tomentosa Original  
      specimen: Panama Canal, Panama [USNM 1521275] Leptuca tomentosa Original  
      specimen: Panama City, Panama [USNM 138840] Leptuca tomentosa Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23051] Leptuca tomentosa Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376360] Leptuca tomentosa Original  
      specimen: Puntarenas, Costa Rica [USNM 137411] Leptuca tomentosa Original  
      specimen: Puntarenas, Costa Rica [USNM 137412] Leptuca tomentosa Original  
      specimen: Puntarenas, Costa Rica [USNM 79406] Leptuca tomentosa Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [USM 802] Leptuca tomentosa Original  
      specimen: Rio Villamil, Guayas, Ecuador [UNI 750] Leptuca tomentosa Original  
      specimen: Rio Villamil, Guayas, Ecuador [UNI P0217] Leptuca tomentosa Original  
      specimen: Rio Villamil, Guayas, Ecuador [UNI P0218] Leptuca tomentosa Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 804] Leptuca tomentosa Original  
 
Leptuca umbratila   text p. 2-3, 6, 9, 13, 31-34, 36; Figure 28; Supplement p. 9 citation: von Prahl (1982)Uca (Boboruca) thayeri umbratila Leptuca umbratila Computed  
      specimen: Diablo, Panama City, Panama [NCHZOOL 13579] Leptuca umbratila Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 805] Leptuca umbratila Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 807] Leptuca umbratila Original  
      specimen: Nicoya Peninsula, Guanacaste, Costa Rica [USNM 138132] Leptuca umbratila Original  
      specimen: Nicoya Peninsula, Guanacaste, Costa Rica [USNM 138133] Leptuca umbratila Original  
      specimen: Panama Canal, Panama [USNM 79407] Leptuca umbratila Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376400] Leptuca umbratila Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376407] Leptuca umbratila Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 806] Leptuca umbratila Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI P0027] Leptuca umbratila Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376402] Leptuca umbratila Original  
 
Minuca   text p. 1, 4, 6, 8-9, 12, 18, 32, 35-36 citation: Shih et al. (2016)Minuca Genus Minuca Computed in part
      citation: Thurman et al. (2023) TBD Computed  
      location: Ecuador Genus Minuca Original  
 
Minuca aff. ecuadoriensis   text p. 10, 20, 32-34; Supplement p. 5 citation: Crane (1975)Uca (Minuca) vocator ecuadoriensis Minuca ecuadoriensis Computed  
      citation: von Hagen (1968)Uca lanigera Minuca ecuadoriensis Computed  
      specimen: Isla Santay, Rio Guayas, Guayas, Ecuador [UNI 746] ? Original  
      specimen: Rio Guayas, Guayas, Ecuador [UNI P0224] ? Original  
 
Minuca aff. zacae   text p. 10, 32-34 specimen: Rio Verde, Esmeraldas, Ecuador [UNI P0040] ? Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI P0041] ? Original  
 
Minuca argillicola   text p. 2-3, 6, 9, 12, 18, 33-34, 37; Figure 9; Supplement p. 4 citation: Landstorfer & Schubart (2010)Uca argillicola Minuca argillicola Computed  
      citation: Rosenberg (2020)Minuca argillicola Minuca argillicola Computed  
      citation: von Prahl (1982)Uca (Celuca) argillicola Minuca argillicola Computed  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 827] Minuca argillicola Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137400] Minuca argillicola Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137401] Minuca argillicola Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376428] Minuca argillicola Original  
      specimen: Isla Portete, Esmeraldas, Ecuador [UNI P0299] Minuca argillicola Original  
      specimen: Isla Portete, Esmeraldas, Ecuador [UNI P0300] Minuca argillicola Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 786] Minuca argillicola Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 824] Minuca argillicola Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI P0016] Minuca argillicola Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 808] Minuca argillicola Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 825] Minuca argillicola Original  
      specimen: Puerto El Morro, Guayas, Ecuador [SMF 34737] Minuca argillicola Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376418] Minuca argillicola Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 823] Minuca argillicola Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 828] Minuca argillicola Original  
      specimen: Rio Esmeraldas, Esmeraldas, Ecuador [UNI 829] Minuca argillicola Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 826] Minuca argillicola Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 855] Minuca argillicola Original  
      specimen: Santo Domingo, Puntarenas, Costa Rica [USNM 19441] Minuca argillicola Original  
 
Minuca brevifrons   text p. 6, 10, 12, 18-19, 33-34, 36; Figure 10; Supplement p. 4 citation: Ramos-Veliz et al. (2022)Minuca brevifrons Minuca brevifrons Computed in part
      specimen: Bahia Cana Blanca, Puntarenas, Costa Rica [SMF 34738] Minuca brevifrons Original  
      specimen: Baja California, Mexico [RMNH.CRUS.D7573] Minuca brevifrons Original  
      specimen: Daule, Esmeraldas, Ecuador [UNI P0302] Minuca brevifrons Original  
      specimen: El Salvador [RMNH.CRUS.D9500] Minuca brevifrons Original  
      specimen: Estero Salado, Guayaquil, Guayas, Ecuador [USNM 70867] Minuca brevifrons Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376435] Minuca brevifrons Original  
      specimen: Guerrero, Mexico [AMNH_IZC 376437] Minuca brevifrons Original  
      specimen: Islas Negritos, Puntarenas, Costa Rica [USNM 138484] Minuca brevifrons Original  
      specimen: Jalisco, Mexico [AMNH_IZC 376439] Minuca brevifrons Original  
      specimen: La Propicia, Esmeraldas, Ecuador [UNI 832] Minuca brevifrons Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 856] Minuca brevifrons Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI P0009] Minuca brevifrons Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 857] Minuca brevifrons Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 858] Minuca brevifrons Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376438] Minuca brevifrons Original  
      specimen: Puente Valdivia, Santa Elena, Ecuador [UNI 842] Minuca brevifrons Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376431] Minuca brevifrons Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 809] Minuca brevifrons Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 859] Minuca brevifrons Original  
      specimen: San Juanillo, Guanacaste, Costa Rica [ZRC 2012.0126] Minuca brevifrons Original  
 
Minuca ecuadoriensis   text p. 2-3, 6, 10, 12, 19-20, 32-34, 36; Figure 11; Supplement p. 4-5 citation: Crane (1975)Uca (Minuca) vocator ecuadoriensis Minuca ecuadoriensis Computed  
      citation: Maccagno (1928)Uca ecuadoriensis Minuca ecuadoriensis Computed  
      citation: von Hagen (1968)Uca lanigera Minuca ecuadoriensis Computed  
      specimen: Baja California, Mexico [AMNH_IZC 376466] Minuca ecuadoriensis Original  
      specimen: Colima, Mexico [AMNH_IZC 376477] Minuca ecuadoriensis Original  
      specimen: Estero Salado, Guayaquil, Guayas, Ecuador [USNM 70867] Minuca ecuadoriensis Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376462] Minuca ecuadoriensis Original  
      specimen: Guerrero, Mexico [AMNH_IZC 376478] Minuca ecuadoriensis Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 811] Minuca ecuadoriensis Original  
      specimen: Mompiche, Esmeraldas, Ecuador [UNI 813] Minuca ecuadoriensis Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 815] Minuca ecuadoriensis Original  
      specimen: Nayarit, Mexico [AMNH_IZC 376461] Minuca ecuadoriensis Original  
      specimen: Nayarit, Mexico [USNM 97756] Minuca ecuadoriensis Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376469] Minuca ecuadoriensis Original  
      specimen: Playa Don Juan, Manabí, Ecuador [UNI 747] Minuca ecuadoriensis Original  
      specimen: Puenta Sitio Nuevo, Santa Elena, Ecuador [UNI 817] Minuca ecuadoriensis Original  
      specimen: Puenta Sitio Nuevo, Santa Elena, Ecuador [UNI P0132] Minuca ecuadoriensis Original  
      specimen: Puente Atravezando, Santa Elena, Ecuador [UNI 816] Minuca ecuadoriensis Original  
      specimen: Puerto El Morro, Guayas, Ecuador [SMF 34740] Minuca ecuadoriensis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23049] Minuca ecuadoriensis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23050] Minuca ecuadoriensis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376487] Minuca ecuadoriensis Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 812] Minuca ecuadoriensis Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 810] Minuca ecuadoriensis Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 376486] Minuca ecuadoriensis Original  
      specimen: Santa Marta, Esmeraldas, Ecuador [UNI 814] Minuca ecuadoriensis Original  
 
Minuca galapagensis   text p. 2-3, 6, 12, 20, 33-34, 37; Figure 12; Supplement p. 5 citation: Capparelli et al. (2021) TBD Computed  
      citation: Rathbun (1918)Uca galapagensis Minuca galapagensis Computed  
      citation: Rosenberg (2020)Minuca galapagensis Minuca galapagensis Computed  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 841] Minuca galapagensis Original  
      specimen: Guayas, Ecuador [USNM 98046] Minuca galapagensis Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 843] Minuca galapagensis Original  
      specimen: Manglaraltos, Santa Elena, Ecuador [UNI 845] Minuca galapagensis Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 830] Minuca galapagensis Original  
      specimen: Muisne, Esmeraldas, Ecuador [UNI 834] Minuca galapagensis Original  
      specimen: Playa Don Juan, Manabí, Ecuador [UNI 748] Minuca galapagensis Original  
      specimen: Puenta Las Javitas, Santa Elena, Ecuador [UNI 837] Minuca galapagensis Original  
      specimen: Puenta Sitio Nuevo, Santa Elena, Ecuador [UNI 838] Minuca galapagensis Original  
      specimen: Puente Atravezando, Santa Elena, Ecuador [UNI 844] Minuca galapagensis Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138536] Minuca galapagensis Original  
      specimen: Puerto El Morro, Guayas, Ecuador [SMF 34741] Minuca galapagensis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D18797] Minuca galapagensis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23048] Minuca galapagensis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D45382] Minuca galapagensis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376489] Minuca galapagensis Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 836] Minuca galapagensis Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 835] Minuca galapagensis Original  
      specimen: Rio Villamil, Guayas, Ecuador [UNI 839] Minuca galapagensis Original  
      specimen: San Clemente, Manabí, Ecuador [UNI P0208] Minuca galapagensis Original  
      specimen: San Clemente, Manabí, Ecuador [UNI P0209] Minuca galapagensis Original  
      specimen: Santa Cruz Island, Galápagos Islands [UNI 831] Minuca galapagensis Original  
      specimen: Santa Cruz Island, Galápagos Islands [UNI P0068] Minuca galapagensis Original  
      specimen: Santa Cruz Island, Galápagos Islands [USNM 22319] Minuca galapagensis Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 833] Minuca galapagensis Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI 846] Minuca galapagensis Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI P0133] Minuca galapagensis Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI P0168] Minuca galapagensis Original  
      specimen: Tortuga Bay, Santa Cruz Island, Galápagos Islands [UNI 840] Minuca galapagensis Original  
      specimen: Tumbes Province, Tumbes Region, Peru [SMF 13151] Minuca galapagensis Original  
 
Minuca herradurensis   text p. 3, 6, 20, 33; Supplement p. 5 citation: Crane (1975)Uca (Minuca) [galapagensis] galapagensis herradurensis Minuca herradurensis Computed in part
      citation: Rathbun (1918)Uca macrodactylus Minuca herradurensis Computed in part; in part
      citation: Rosenberg (2020)Minuca herradurensis Minuca herradurensis Computed  
      specimen: Honduras [AMNH_IZC 376584] Minuca herradurensis Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376506] Minuca herradurensis Original  
      specimen: Panama City, Panama [USNM 138542] Minuca herradurensis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376500] Minuca herradurensis Original  
      specimen: Puntarenas, Costa Rica [USNM 123790] Minuca herradurensis Original  
      specimen: Salina Cruz, Oaxaca, Mexico [AMNH_IZC 376615] Minuca herradurensis Original  
 
Minuca osa   text p. 6, 10, 12, 20-21, 33-34, 36; Figure 13; Supplement p. 5-6 citation: Landstorfer & Schubart (2010)Uca osa Minuca osa Computed  
      citation: Lombardo-Gonzalez (2022) TBD    
      citation: Rosenberg (2020)Minuca osa Minuca osa Computed  
      specimen: Golfo Dolce, Puntarenas, Costa Rica [RMNH.CRUS.D53098] Minuca brevifrons Original  
      specimen: Golfo Dolce, Puntarenas, Costa Rica [RMNH.CRUS.D53126] Minuca brevifrons Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 820] Minuca osa Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI 822] Minuca osa Original  
      specimen: La Tola, Esmeraldas, Ecuador [UNI P0303] Minuca brevifrons Original  
      specimen: Puente Atravezando, Santa Elena, Ecuador [UNI 819] Minuca osa Original  
      specimen: Rio Agua Clara, Esmeraldas, Ecuador [UNI 818] Minuca osa Original  
      specimen: Rio Cadente, Santa Elena, Ecuador [UNI 821] Minuca osa Original  
      specimen: Rio Verde, Esmeraldas, Ecuador [UNI 860] Minuca osa Original  
      specimen: Simon Bolivar, Santa Elena, Ecuador [UNI P0123] Minuca brevifrons Original  
 
Minuca panema   text p. 33 citation: Thurman et al. (2021) TBD Computed  
      citation: Thurman et al. (2023) TBD Computed  
 
Minuca pygmaea   text p. 2, 6, 33 citation: Rosenberg (2020)Leptuca pygmaea Leptuca pygmaea Computed Authors use a different genus for this species in the text and supplemental information
      citation: von Prahl (1982)Uca (Minuca) pygmaea Leptuca pygmaea Computed Authors use a different genus for this species in the text and supplemental information
      specimen: Bahia de Buenaventura, Valle del Cauca, Colombia [USNM 18646] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137419] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137420] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
      specimen: Golfito, Puntarenas, Costa Rica [USNM 79402] Leptuca pygmaea Original Authors use a different genus for this species in the text and supplemental information
 
Minuca pygmeae   text p. 3 citation: von Prahl (1982)Uca (Minuca) pygmaea Leptuca pygmaea Computed typo of species name
 
Minuca virens   text p. 33 citation: Salmon & Atsaides (1968)Uca virens Minuca virens Computed  
      citation: Thurman et al. (2018) TBD Computed  
 
Minuca zacae   text p. 6, 10, 33-34; Supplement p. 6 citation: Crane (1941)Uca zacae Minuca zacae Computed  
      citation: Rosenberg (2020)Minuca zacae Minuca zacae Computed  
      specimen: Colima, Mexico [AMNH_IZC 376370] Minuca zacae Original  
      specimen: Corinto, Nicaragua [USNM 79408] Minuca zacae Original  
      specimen: El Salvador [SMF 2104a] Minuca zacae Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137426] Minuca zacae Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 137427] Minuca zacae Original  
      specimen: Golfo Dolce, Puntarenas, Costa Rica [USNM 138646] Minuca zacae Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376728] Minuca zacae Original  
      specimen: Guerrero, Mexico [AMNH_IZC 376723] Minuca zacae Original  
      specimen: Islas Negritos, Puntarenas, Costa Rica [USNM 1521269] Minuca zacae Original  
      specimen: Los Blancos, La Paz, El Salvador [ZMA.CRUS.D100.365] Minuca zacae Original  
      specimen: Nayarit, Mexico [AMNH_IZC 376716] Minuca zacae Original  
      specimen: Oaxaca, Mexico [AMNH_IZC 376715] Minuca zacae Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376718] Minuca zacae Original  
 
Petruca   text p. 1, 4, 6-8, 11, 16, 32, 36 citation: Shih et al. (2016)Petruca TBD Computed  
      location: Ecuador Genus Petruca Original  
 
Petruca panamensis   text p. 2-3, 6-7, 11, 16-18, 34-35, 37; Figure 8; Supplement p. 4 citation: Rathbun (1918)Uca panamensis Petruca panamensis Computed  
      citation: Shih et al. (2015) TBD Computed  
      specimen: Costa Rica [ZRC] Petruca panamensis Original  
      specimen: Golfito, Puntarenas, Costa Rica [USNM 138625] Petruca panamensis Original  
      specimen: Gorgona Island, Cauca, Colombia [USNM 138629] Petruca panamensis Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13581] Petruca panamensis Original  
      specimen: Isla Naos, Panama [USNM 1294205] Petruca panamensis Original  
      specimen: Playa Camarones, Esmeraldas, Ecuador [UNI 752] Petruca panamensis Original  
      specimen: Playa Camarones, Esmeraldas, Ecuador [UNI P0293] Petruca panamensis Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376770] Petruca panamensis Original  
      specimen: Salinas, Santa Elena, Ecuador [USNM 70989] Petruca panamensis Original  
      specimen: San Juanillo, Guanacaste, Costa Rica [NCHUZOOL 14753] Petruca panamensis Original  
      specimen: San Juanillo, Guanacaste, Costa Rica [ZRC 2012.0126] Petruca panamensis Original  
 
Tubuca alcocki   text p. 33, 35 citation: Shih et al. (2018)Tubuca alcocki Tubuca alcocki Computed in part
 
Tubuca urvillei   text p. 35 citation: Shih et al. (2018)Tubuca urvillei Tubuca urvillei Computed in part
 
Uca   text p. 1, 4-5, 7-8, 11, 13, 32, 36 citation: Shih et al. (2016)Uca TBD Computed in part
      location: Ecuador Genus Uca Original  
 
Uca heteropleura   text p. 2-3, 5, 7, 11-13, 34, 36; Figure 2; Supplement p. 3 citation: von Hagen (1968)Uca heteropleura Uca heteropleura Computed  
      specimen: Posorja, Guayas, Ecuador [UNI 743] Uca heteropleura Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138550] Uca heteropleura Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138577] Uca heteropleura Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23059] Uca heteropleura Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376846] Uca heteropleura Original  
      specimen: Salinas, Santa Elena, Ecuador [UNI P0240] Uca heteropleura Original  
      specimen: San Clemente, Manabí, Ecuador [UNI 744] Uca heteropleura Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 753] Uca heteropleura Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 755] Uca heteropleura Original  
 
Uca inaequalis   text p. 2 citation: Rathbun (1935)Uca inaequalis Leptuca inaequalis Computed  
 
Uca insignis   text p. 2-3, 5-7, 11-14, 34, 36; Figure 3; Supplement p. 3 citation: Rathbun (1918)Uca insignis Uca insignis Computed in part
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138577] Uca insignis Original  
      specimen: Puerto El Morro, Guayas, Ecuador [SMF 34743] Uca insignis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23063] Uca insignis Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [USNM 1521395] Uca insignis Original  
 
Uca intermedia   text p. 2-3, 5-7, 11, 14, 33-34, 36; Figure 4; Supplement p. 3 citation: Rosenberg (2020)Uca intermedia Uca intermedia Computed  
      citation: von Prahl & Toro (1985)Uca (Uca) intermedia Uca intermedia Computed  
      specimen: Bahia de Buenaventura, Valle del Cauca, Colombia [RMNH.CRUS.D35788] Uca intermedia Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 754] Uca intermedia Original  
      specimen: San Clemente, Manabí, Ecuador [UNI 881] Uca intermedia Original  
      specimen: [RMNH.CRUS.D023063] Uca intermedia Original  
 
Uca ornata   text p. 2-3, 5-7, 14-15, 33, 36; Figure 5; Supplement p. 3 citation: Crane (1975)Uca (Uca) ornata Uca ornata Computed in part
      citation: von Hagen (1968)Uca pizarri Uca ornata Computed  
      specimen: Guayaquil, Guayas, Ecuador [RMNH.CRUS.D23062] Uca ornata Original  
      specimen: Panama City, Panama [USNM 134616] Uca ornata Original  
      specimen: Panama City, Panama [USNM 138614] Uca ornata Original  
      specimen: Panama City, Panama [USNM 138615] Uca ornata Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23061] Uca ornata Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376865] Uca ornata Original  
 
Uca princeps   text p. 2-3, 5, 7, 11, 15-16, 34, 36; Figure 6; Supplement p. 3 citation: Rathbun (1918)Uca princeps Uca princeps Computed in part
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI 854] Uca princeps Original  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI P0252] Uca princeps Original  
      specimen: Baja California, Mexico [AMNH_IZC 376866] Uca princeps Original  
      specimen: Colima, Mexico [AMNH_IZC 376877] Uca princeps Original  
      specimen: Guaymas, Sonora, Mexico [RMNH.CRUS.D.7575] Uca princeps Original  
      specimen: Gulf of California, Mexico [RMNH D250] Uca princeps Original  
      specimen: Nayarit, Mexico [AMNH_IZC 376879] Uca princeps Original  
      specimen: Posorja, Guayas, Ecuador [UNI 741] Uca princeps Original  
      specimen: Puenta Sitio Nuevo, Santa Elena, Ecuador [UNI 757] Uca princeps Original  
      specimen: Puento Carnero, Santa Elena, Ecuador [UNI 756] Uca princeps Original  
      specimen: Puento Carnero, Santa Elena, Ecuador [UNI P0197] Uca princeps Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138633] Uca princeps Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138634] Uca princeps Original  
      specimen: Puerto El Morro, Guayas, Ecuador [UNI P0195] Uca princeps Original  
      specimen: Puerto El Morro, Guayas, Ecuador [UNI P0199] Uca princeps Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D.23060] Uca princeps Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376867] Uca princeps Original  
      specimen: Salinas, Santa Elena, Ecuador [UNI P0241] Uca princeps Original  
      specimen: San Clemente, Manabí, Ecuador [UNI 745] Uca princeps Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 758] Uca princeps Original  
      specimen: San Lorenzo, Honduras [AMNH_IZC 377881] Uca princeps Original  
      specimen: Tumbes Province, Tumbes Region, Peru [SMF 13164] Uca princeps Original  
 
Uca stylifera   text p. 2-3, 5, 7, 11, 16-17, 34, 36; Figure 7; Supplement p. 3-4 citation: Rathbun (1918)Uca stylifera Uca stylifera Computed in part
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI 742] Uca stylifera Original  
      specimen: Bahia de Caraquez, Manabí, Ecuador [UNI P0260] Uca stylifera Original  
      specimen: Balboa, Panama City, Panama [USNM 138836] Uca stylifera Original  
      specimen: Base Naval Vasco Nuñez de Balboa, Panama [NCHUZOOL 13578] Uca stylifera Original  
      specimen: El Rompido, Esmeraldas, Ecuador [UNI 759] Uca stylifera Original  
      specimen: Guanacaste, Costa Rica [AMNH_IZC 376895] Uca stylifera Original  
      specimen: Isla Portete, Esmeraldas, Ecuador [UNI P0286] Uca stylifera Original  
      specimen: Puerto Bolivar, El Oro, Ecuador [USNM 138837] Uca stylifera Original  
      specimen: Puerto Pizarro, Tumbes Province, Tumbes Region, Peru [RMNH.CRUS.D23058] Uca stylifera Original  
      specimen: Puntarenas, Costa Rica [AMNH_IZC 376889] Uca stylifera Original  
      specimen: Puntarenas, Costa Rica [USNM 32325] Uca stylifera Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 760] Uca stylifera Original  
      specimen: San Jose de Chamanga, Esmeraldas, Ecuador [UNI 761] Uca stylifera Original  
 
Xeruca formosensis   text p. 35 citation: Shih (2015) TBD Computed