von Hagen, H.-O. (1968) Studien an peruanischen Winkerkrabben (Uca). Zoologische Jahrbücher. Abteilung für Systematik, Ökologie und Geographie der Tiere 95(2):395–468.
Purple fiddler crabs indicate locations where this species is found according to the scientific record; blue fiddler crabs surrounded by a dashed circle indicate the same but for larger, general regions; green fiddler crabs indicate “research grade” observations imported from iNaturalist; yellow circles with a question mark indicate questionable records of this species from the scientific record; and red circles with an x indicate false or mistaken records from the scientific record.